Because caged birds become disoriented when the sky is overcast, Kramer hypothesized that birds orient themselves according to the Sun's position. | In one set of experiments, Kramer placed the box containing food at the same point of the compass each time he put food boxes in the birds’ environment. | Kramer demonstrated that an internal biological clock allows starlings to compensate for the Sun's movement. | After several studies, Kramer surmised that an internal biological clock allows some species of birds to navigate at night. | The role of environmental cues in birds' navigation is clear, for on overcast days, birds use objects besides the Sun to orient themselves. | Kramer showed that night-migrating birds use the sky to navigate by the stars. |